Dung Beetles of Manyoni Private Game Reserve

Kheper nigroaeneus (Scarabaeus nigroaeneus) and Garreta wahlbergi (Garreta nitens)

Chalconotus convexus

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Copris amyntor

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Copris bootes

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Copris denticulatus

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Digitonthophagus gazella

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Garreta nitens

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Heliocopris hamadryas

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Heliocopris neptunus

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Kheper nigroaeneus

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Kheper subaeneus

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Onitis fulgidus

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Scarabaeus deludens

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Sisyphus sordidus

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