File:Alouatta guariba (Brown Howler Monkey).jpg

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Description	English: Brown Howler Monkey (Alouatta guariba) in Feliciano Miguel Abdala Reserve, Caratinga, Brazil.
Date	21 October 2011, 15:20:47
Source	Flickr
Author	Bart van Dorp

Licensing edit

This image was originally posted to Flickr by Bart vanDorp at https://www.flickr.com/photos/39351863@N06/6282812706. It was reviewed on 19 July 2013 by FlickreviewR and was confirmed to be licensed under the terms of the cc-by-2.0.

19 July 2013

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	Date/Time	Thumbnail	Dimensions	User	Comment
current	18:06, 19 July 2013		1,446 × 932 (1.16 MB)	Miguelrangeljr (talk \| contribs)	Watermark removed (cropped).
	17:31, 19 July 2013		1,772 × 1,184 (908 KB)	Miguelrangeljr (talk \| contribs)	User created page with UploadWizard

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Usage on en.wikipedia.org
- Feliciano Miguel Abdala Private Natural Heritage Reserve
Usage on fr.wikipedia.org
- Liste des 25 primates les plus menacés au monde

Metadata

This file contains additional information such as Exif metadata which may have been added by the digital camera, scanner, or software program used to create or digitize it. If the file has been modified from its original state, some details such as the timestamp may not fully reflect those of the original file. The timestamp is only as accurate as the clock in the camera, and it may be completely wrong.

Camera manufacturer	Canon
Camera model	Canon EOS-1D Mark IV
Author	Bart van Dorp
Exposure time	1/300 sec (0.0033333333333333)
F-number	f/7.1
ISO speed rating	640
Date and time of data generation	15:20, 21 October 2011
Lens focal length	300 mm
Image title	Also know as: Brown howling monkey, Bugio, guariba, macaco-uivador.Eleé famoso por seu grito, que pode ser ouvido em toda a mata, e pela presença de pêlos mais compridos nos lados da face formando uma espécie de barba.As their name suggests, vocal communication forms an important part of their social behavior. They have an enlarged basihyal or hyoid bone which helps them make their loud vocalizations. Group males generally call at dawn and dusk as well as interspersed times throughout the day. The main vocals consist of loud, deep guttural growls or "howls." Howler monkeys are widely considered to be the loudest land animal. According to Guinness Book of World Records, their vocalizations can be heard clearly for 3 miles (4.8 km). It is hypothesized that the
Short title	Barbado / Brown howler (Alouatta guariba)
Width	1,772 px
Height	1,184 px
Bits per component	8 8 8
Pixel composition	RGB
Orientation	Normal
Number of components	3
Horizontal resolution	150 dpi
Vertical resolution	150 dpi
Software used	Adobe Photoshop CS5 Windows
File change date and time	14:50, 19 July 2013
Exposure Program	Manual
Exif version	2.3
Date and time of digitizing	15:20, 21 October 2011
APEX shutter speed	8.2288192128085
APEX aperture	5.6556380823768
APEX exposure bias	0
Maximum land aperture	5 APEX (f/5.66)
Subject distance	12.4 meters
Metering mode	Pattern
Flash	Flash fired, compulsory flash firing
DateTimeOriginal subseconds	46
DateTimeDigitized subseconds	46
Color space	sRGB
Focal plane X resolution	3,795.3488372093
Focal plane Y resolution	3,904.306122449
Focal plane resolution unit	inches
Custom image processing	Normal process
Exposure mode	Manual exposure
White balance	Auto white balance
Scene capture type	Standard
Serial number of camera	620601784
Lens used	EF70-300mm f/4-5.6L IS USM
Date metadata was last modified	11:50, 19 July 2013
Unique ID of original document	xmp.did:350523AA9BF0E211B922FB9888A8761A
Keywords	brazil macaco-uivador MG Atlantic Forest Bart van Dorp Caratinga Barbado Mata Atlântica brasil Reserva Particular do Património Natural Feliciano Miguel Abdala guariba RPPN Feliciano Miguel Abdala Bugio Bart vanDorp Brown howler Brown howling monkey primatas
IIM version	2