In camp at Kill Devil Hills, the Wright Brothers endured weeks of delays caused by broken propeller shafts during engine tests. After the shafts were replaced (requiring two trips back to Dayton), Wilbur won a coin toss and made a three-second flight attempt on December 14, 1903, stalling after takeoff and causing minor damage to the Flyer. (Because December 13, 1903, was a Sunday, the brothers did not make any attempts that day, even though the weather was good.) In a message to their family, Wilbur referred to the trial as having "only partial success", stating "the power is ample, and but for a trifling error due to lack of experience with this machine and this method of starting, the machine would undoubtedly have flown beautifully."

Following repairs, the Wrights finally took to the air on December 17, 1903, making two flights each from level ground into a freezing headwind gusting to 27 miles per hour (43 km/h). The first flight, by Orville at 10:35 am, of 120 feet (37 m) in 12 seconds, at a speed of only 6.8 miles per hour (10.9 km/h) over the ground, was recorded in a famous photograph. The next two flights covered approximately 175 and 200 feet (53 and 61 m), by Wilbur and Orville respectively. Their altitude was about 10 feet (3.0 m) above the ground. The following is Orville Wright's account of the final flight of the day:

Wilbur started the fourth and last flight at just about 12 o'clock. The first few hundred feet were up and down, as before, but by the time three hundred ft had been covered, the machine was under much better control. The course for the next four or five hundred feet had but little undulation. However, when out about eight hundred feet the machine began pitching again, and, in one of its darts downward, struck the ground. The distance over the ground was measured to be 852 feet; the time of the flight was 59 seconds. The frame supporting the front rudder was badly broken, but the main part of the machine was not injured at all. We estimated that the machine could be put in condition for flight again in about a day or two.

Five people witnessed the flights: Adam Etheridge, John T. Daniels (who snapped the famous "first flight" photo using Orville's pre-positioned camera) and Will Dough, all of the U.S. government coastal lifesaving crew; area businessman W.C. Brinkley; and Johnny Moore, a teenaged boy who lived in the area. After the men hauled the Flyer back from its fourth flight, a powerful gust of wind flipped it over several times, despite the crew's attempt to hold it down. Severely damaged, the airplane never flew again. The brothers shipped it home, and years later Orville restored it, lending it to several U.S. locations for display, then to a British museum, before it was finally installed in the Smithsonian Institution in Washington, D.C. in 1948, its current residence.

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