Smith System

English: The Smith System is a fern classification system that was co-developed by Alan R. Smith, Research Botanist at the University of California, Berkeley, in 2006. It is often referenced as Smith et al. 2006 or Smith & al. 2006.

Phylogenies edit

Pteridophyta phylogeny (es)
Monilophyte phylogeny (es)
Leptosproangiates phylogeny (es)
Polypodiales phylogeny (es)

Circumscription of living taxa sensu Smith et al. 2006 ^[1] corrected in Cyatheaceae sensu Korall et al. 2007^[2]

Psilotopsida
- Psilotales
  - Psilotaceae
    - Psilotum
    - Tmesipteris
- Ophioglossales
  - Ophioglossaceae
    - Botrychium (included Sceptridium, Botrypus, Japanobotrychium)
    - Helminthostachys
    - Mankyua (from Cheju island, Korea)
    - Ophioglossum (incl. Cheiroglossa, Ophioderma)
Equisetopsida
- Equisetales
- Sphenophyllales
Marattiopsida
- Marattiales
  - Marattiaceae
    - Angiopteris (incl. Archangiopteris)
    - Christensenia
    - Danaea
    - Marattia
Polypodiopsida (= Filicopsida, = "leptosporangiate ferns")
- Osmundales
  - Osmundaceae
    - Leptopteris
    - Osmunda
    - Osmundastrum
    - Todea
- Hymenophyllales
  - Hymenophyllaceae
    - Hymenophyllum (including Cardiomanes, Hymenoglossum, Rosenstockia, Serpyllopsis)
    - Abrodictyum
    - Callistopteris
    - Cephalomanes
    - Crepidomanes
    - Didymoglossum
    - Polyphlebium
    - Trichomanes sensu stricto (this is NOT Trichomanes as usually seen)
    - Vandenboschia
- Gleicheniales
  - Gleicheniaceae
    - Dicranopteris
    - Diplopterygium
    - Gleichenella
    - Gleichenia
    - Sticherus
    - Stromatopteris
  - Dipteridaceae
    - Cheiropleuria
    - Dipteris
  - Matoniaceae
    - Matonia
    - Phanerosorus
- Schizaeales
  - Lygodiaceae
    - Lygodium
  - Schizaeaceae
    - Schizaea
    - Actinostachys
  - Anemiaceae
    - Anemia
- Salviniales
  - Marsileaceae
    - Marsilea
    - Pilularia
    - Regnellidium
  - Salviniaceae
    - Salvinia
    - Azolla
- Cyatheales ("tree ferns clade").
  - Thyrsopteridaceae
    - Thyrsopteris elegans
  - Loxomataceae
    - Loxoma
    - Loxsomopsis
  - Culcitaceae
    - Culcita
  - Plagiogyriaceae
    - Plagiogyria
  - Cibotiaceae
    - Cibotium
  - Cyatheaceae
    - Alsophila (including Gymnosphaera, Nephelea)
    - Cyathea (incl. Hymenophyllopsis, Cnemidaria, Hemitelia, Trichipteris)
    - Sphaeropteris (incl. Schizocaena, Fourniera).
  - Dicksoniaceae
    - Calochlaena
    - Dicksonia
    - Lophosoria
  - Metaxyaceae
    - Metaxya
- Polypodiales
  - Lindsaeaceae
    - Cystodium
    - Lindsaea
    - Lonchitis
    - Odontosoria
    - Ormoloma
    - Sphenomeris
    - Tapeinidium
    - Xyropteris
  - Saccolomataceae
    - Saccoloma
  - Dennstaedtiaceae
    - Blotiella
    - Coptodipteris
    - Dennstaedtia (incl. Costaricia)
    - Histiopteris
    - Hypolepis
    - Leptolepia
    - Microlepia
    - Monachosorum
    - Oenotrichia sensu stricto
    - Paesia
    - Pteridium
  - Pteridaceae 50 genera, some of them:
    - Acrostichum
    - Actiniopteris
    - Adiantopsis
    - Adiantum
    - Aleuritopteris
    - Ananthacorus (vittarioide)
    - Anetium (vittarioide)
    - Anogramma
    - Antrophyum (vittarioide)
    - Argyrochosma
    - Aspidotis
    - Astrolepis
    - Austrogramme
    - Bommeria
    - Cassebeera
    - Ceratopteris
    - Cerosora
    - Cheilanthes
    - Cheiloplecton
    - Coniogramme
    - Cosentinia (Nakazato y Gastony 2001)
    - Cryptogramma
    - Doryopteris
    - Eriosorus
    - Haplopteris (vittarioide)
    - Hecistopteris (vittarioide)
    - Hemionitis
    - Holcochlaena
    - Jamesonia
    - Llavea
    - Mildella
    - Monogramma (vittarioide)
    - Nephopteris
    - Neurocallis
    - Notholaena
    - Ochropteris
    - Onychium
    - Paraceterach
    - Parahemionitis
    - Pellaea
    - Pentagramma
    - Pityrogramma
    - Platyloma
    - Platyzoma
    - Polytaenium (vittarioide)
    - Pteris (incl. Afropteris, Anopteris)
    - Pterozonium
    - Radiovittaria (vittarioide)
    - Rheopteris (vittarioide)
    - Scoliosorus (vittarioide)
    - Syngramma
    - Taenitis
    - Trachypteris
    - Vittaria (vittarioide)
  - Aspleniaceae:
    - Asplenium (included Camptosorus, Loxoscaphe, Diellia, Pleurosorus, Phyllitis, Ceterach, Thamnopteris, and others, and probably also included Antigramma, Holodictyum,Schoffneria,Sinephropteris)
    - Hymenasplenium
  - Woodsiaceae
    - Athyrium
    - Diplazium (incl. Callipteris,Monomelangium)
    - Acystopteris
    - Cheilanthopsis
    - Cornopteris
    - Cystopteris
    - Deparia (incl. Lunathyrium, Dryoathyrium, Athyriopsis, Dictyodroma)
    - Diplaziopsis
    - Gymnocarpium (incl.Currania)
    - Hemidictyum
    - Homalosorus
    - Protowoodsia
    - Pseudocystopteris
    - Rhachidosorus
    - Woodsia (incl. Hymenocystis)
  - Thelypteridaceae
    - Cyclosorus (incl. Ampelopteris, Amphineuron, Chingia, Christella, Cyclogramma, Cyclosorus sensu stricto, Glaphyropteridopsis, Goniopteris, Meniscium, Menisorus, Mesophlebion, Pelazoneuron, Plesioneuron, Pneumatopteris, Pronephrium, Pseudocyclosorus, Sphaerostephanos, Stegnogramma, Steiropteris, Trigonospora)
    - Macrothelypteris
    - Phegopteris
    - Pseudophegopteris
    - Thelypteris (incl. Amauropelta, Coryphopteris, Metathelypteris, Oreopteris, Parathelypteris).
  - Blechnaceae
    - Blechnum sensu lato
    - Brainea
    - Doodia
    - Pteridoblechnum
    - Sadleria
    - Salpichlaena
    - Steenisioblechnum
    - Stenochlaena
    - Woodwardia (incl. Anchistea,Chieniopteris, Lorinseria).
  - Onocleaceae
    - Matteuccia
    - Onoclea
    - Onocleopsis
    - Pentarhizidium
  - Dryopteridaceae
    - Ctenitis
    - Dryopteris (incl. Nothoperanema)
    - Elaphoglossum (incl. Microstaphyla, Peltapteris)
    - Polystichum (incl. Papuapteris, Plecosorus)
    - Acrophorus
    - Acrorumohra
    - Adenoderris
    - Arachniodes
    - Ataxipteris
    - Bolbitis (incl. Egenolfia)
    - Coveniella
    - Cyclodium
    - Cyrtogonellum
    - Cyrtomidictyum
    - Cyrtomium
    - Didymochlaena
    - Dryopolystichum
    - Dryopsis
    - Hypodematium
    - Lastreopsis
    - Leucostegia
    - Lithostegia
    - Lomagramma
    - Maxonia
    - Megalastrum
    - Oenotrichia p.p.
    - Olfersia
    - Peranema
    - Phanerophlebia
    - Polybotrya
    - Polystichopsis

- - - Revwattsia
    - Rumohra
    - Stenolepia
    - Stigmatopteris
    - Teratophyllum
  - Lomariopsidaceae
    - Cyclopeltis
    - Lomariopsis
    - Nephrolepis
    - Thysanosoria
  - Tectariaceae
    - Tectaria sensu lato (including Amphiblestra, Camptodium, Chlamydogramme, Cionidium, Ctenitopsis, Dictyoxiphium, Fadyenia,Hemigramma, Pleuroderris, Pseudotectaria,Quercifilix, and maybe some of the genera mentioned down here).
    - Aenigmopteris
    - Arthropteris
    - Heterogonium
    - Hypoderris
    - Pleocnemia
    - Psammiosorus
    - Psomiocarpa
    - Pteridrys
    - Triplophyllum
  - Oleandraceae
    - Oleandra
  - Davalliaceae
    - Araiostegia
    - Davallia (incl. Humata,Parasorus,Scyphularia)
    - Davallodes
    - Pachypleuria
  - Polypodiaceae
    - Acrosorus
    - Adenophorus
    - Aglaomorpha (incl. Photinopteris, Merinthosorus, Pseudodrynaria, Holostachyum)
    - Arthromeris
    - Belvisia
    - Calymmodon
    - Campyloneurum
    - Ceradenia
    - Christiopteris
    - Chrysogrammitis
    - Cochlidium
    - Colysis
    - Ctenopteris
    - Dicranoglossum
    - Dictymia
    - Drynaria
    - Enterosora
    - Goniophlebium sensu lato
    - Grammitis
    - Lecanopteris
    - Lellingeria
    - Lemmaphyllum
    - Lepisorus (incl. Platygyria)
    - Leptochilus
    - Loxogramme (incl. Anarthropteris)
    - Melpomene
    - Microgramma
    - Micropolypodium
    - Pyrrosia (incl. Drymoglossum)
    - Scleroglossum
    - Selliguea (incl. Crypsinus, Polypodiopteris)
    - Serpocaulon
    - Synammia
    - Terpsichore
    - Zygophlebia
    - Caobangia
    - Drymotaenium
    - Gymnogrammitis
    - Kontumia
    - Luisma
    - Pleurosoriopsis
    - Podosorus
    - Polypodium
    - Microsorum

References edit

↑ A. R. Smith, K. M. Pryer, E. Schuettpelz, P. Korall, H. Schneider, P. G. Wolf. 2006. "A classification for extant ferns". Taxon 55(3), 705-731 (pdf here)
↑ Korall, P., D.S. Conant, J.S. Metzgar, H. Schneider, K.M. Pryer. 2007. A molecular phylogeny of scaly tree ferns (Cyatheaceae). American Journal of Botany 94: 873-886. (pdf here)

Other classifications edit

List of Classifications used by wikicommons
for Angiosperms:
- Cronquist System
- Strasburger
- APG II in migration to APG III
- APWebsite Classification
for Ferns:
- Smith System
for birds:
- IOC Classification

External Links edit

Smith & al. 2006

[1] A. R. Smith, K. M. Pryer, E. Schuettpelz, P. Korall, H. Schneider, P. G. Wolf. 2006. "A classification for extant ferns". Taxon 55(3), 705-731 (pdf here)

[2] Korall, P., D.S. Conant, J.S. Metzgar, H. Schneider, K.M. Pryer. 2007. A molecular phylogeny of scaly tree ferns (Cyatheaceae). American Journal of Botany 94: 873-886. (pdf here)

[1]

[2]